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He received his bachelors's degree from Washington & Jefferson College, and his Ph.D at the University of the Witwatersrand, South Africa in 1984.
His research focuses on the hominin fossil record, during the Pliocene and early Pleistocene and the reconstruction of phylogenetic relationships through dental morphology. His most important work has been the analysis of dental microwear in order to reconstruct early hominin dietary habits.
Laboratory 1: Anatomical Terminology, General Osteology, and General Arthrology
1.1 Anatomical Terminology
1.2 General Osteology
1.3 General Arthrology
Laboratory 2: The Back
2.1 The Integument
2.2 The Vertebral Column
2.3 Spinal Cord and Spinal Nerves
2.4 Muscles of the Back
Laboratory 3: The Upper Limb
3.1 Bones of the Upper Limb3.2 Joints of the Upper Limb
3.3 Innervation of the Upper Limb
3.4 Muscles of the Upper Limb
3.5 Blood Vessels of the Upper Limb
Laboratory 4: The Lower Limb
4.1 Bones of the Lower Limb4.2 Joints of the Lower Limb
4.3 Innervation of the Lower Limb
4.4 Muscles of the Lower Limb
4.5 Blood Vessels of the Lower Limb
Laboratory 5: The Neck
5.1 Bones and Cartilages of the Neck5.2 Nerves of the Neck
5.3 Muscles of the Neck
5.4 Blood Vessels of the Neck
5.5 Thyroid and Parathyroid Glands
Laboratory 6: The Head
6.1 The Skull6.2 The Dentition
6.3 Muscles of the Head
6.4 Nasal and Oral Cavities
6.5 Blood Vessels of the Head
Laboratory 7: The Brain and Cranial Nerves
7.1 The Brain7.2 Cranial Nerves
Laboratory 8: The Eye and The Ear
8.1 The Eye8.2 The Ear
Laboratory 9: The Thorax
9.1 The Breast9.2 The Thoracic Skeleton
9.3 Skeletal Muscles of the Thorax
9.4 The Thoracic Cavity
9.5 The Mediastinum
9.6 The Respiratory Apparatus
9.7 The Heart
9.8 Lymphatics in the Thorax
9.9 Blood Vessels of the Thorax
9.10 Nerves of the Thorax
Laboratory 10: The Abdomen
10.1 The Abdominal Skeleton10.2 Skeletal Muscles of the Abdomen
10.3 Abdominal Cavity and peritoneum
10.4 Digestive Canal and Organs
10.5 Blood Vessels of the Gut
10.6 Lymphatic Organs and Lymph Drainage
10.7 The Kidneys and Adrenal Glands
10.8 Gonadal Blood Vessels
10.9 Nerves in the Abdomen
Laboratory 11: The Pelvis
11.1 The Pelvic Skeleton11.2 Muscles of the Pelvis
11.3 Peritoneum in the Pelvic Cavity
11.4 Common Pelvic Viscera
11.5 Male Genitalia
11.6 Female Genitalia
11.7 Blood Vessels of the Pelvis
11.8 Nerves of the Pelvis
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